Notes for Chapter 19:
Aquatic Mandibulates (Crustaceans)

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or back to Chapter 18
or ahead to Chapter 20

Chapter 19 Assignment: 389-399, 406-409; RQ-19: 1, 4-9
Next: Ch. 20: 411-427, 432-437; RQ-20: 1-4, 6, 7, 9, 10, 18

Introduction: "Insects" of the Sea


Source of Image

Featured organism: Copepod

More Links: 1 - 2 - 3 - 4 - 5 - 6 - 7 - 8 - 9 - 10 - 11
Images: 1 - 2 - 3

I. Crustacea
    More Links


Features of Crustaceans
        terms: nauplius larva, exoskeleton, molting, copepodite, zoea,
            and megalops larval stages


have 2 prs. antennae and maxillae
most appendages biramous
marine, freshwater, terrestrial
40,000+ species
two pairs of antennae


extensive modification of appendages


hemocoel: blood-filled body space (replaces coelom)
respiration: body surface or gills
circulation: open (
sinuses replace capillaries)
dorsal heart w/ostia
hemocyanin (copper) or hemoglobin (iron)
excretion: antennal (or green) glands
reabsorbs salts, amino acids, water
excretes ammonia (contrast with terrestrial arthropods)
nerves: similar to annelids (paired ventral nerve cord, etc.)
sensory system: statocysts sensory hairs,
compound eyes (ommatidia are the individual elements)
reproduction, growth, and stuff generally dioecious (separate sexes)
(except barnacles and ostracods)
metamorphosis from nauplius, zoeal, cyprid types of larvae
naplius larval type is especially widespread
growth by ecdysis (molting)
molting: allows for growth, sex, etc.
molting is hormonally controlled: x-organ (inhibitor), y-organ (promoter)
feeding: mandibles, maxillae, maxillipeds

II. A Brief Survey of Crustaceans

Get to know underlined groups on hand-out Arthropod classification

Crabs, Shrimps, Lobsters, and Their Relatives

        terms: carapace

        Featured Organism: Sand crabs (Emerita analoga)
        Featured organism: crab, Cancer productus
        Featured Organism: Jonah crab (Cancer borealis)
        Featured Organism: ghost shrimp
        Featured Organism: Copepod
        Featured organism: Barnacle
        Featured organism: Cladoceran (water fleas)

        1) Copepods are central to ocean food webs.

        2) Life cycles reveal that all barnacles are crustaceans - and
            some parasites are barnacles.

        terms: hermaphroditic, nauplius to cypris larval stages


III. Phylogeny and Adaptive Radiation
   More Links: 1 - 2 - 3 - 4 - 5

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or ahead to Chapter 20


This page created 10/14/01 © D.J. Eernisse, Last Modified 10/19/01, Links Last Completely Checked 10/14/01